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- 2. The Cost Of Natural Selection
- 3. In the introduction to The Cost of Natural Selection Haldane writes that it is difficult for
- 4. That is, the problem for the cattle breeder is that keeping only the specimens with the
- 5. Haldane states that this same problem arises with respect to natural selection. Characters that are positively
- 6. Haldane proceeds to define the intensity of selection regarding "juvenile survival" (that is, survival to reproductive
- 7. The proportion for the entire population that die without reproducing is thus 1-S, and this would
- 8. Origin of the term "Haldane's dilemma" Apparently the first use of the term "Haldane's dilemma" was
- 9. Kimura (1960, 1961) has referred to this loss as the substitutional (or evolutional) load, but because
- 10. The Cost Haldane writes, I shall investigate the following case mathematically. A population is in equilibrium
- 11. A sudden change occurs in the environment, for example, pollution by smoke, a change of climate,
- 12. The selective death that must occur for a gene to be substituted was called the cost
- 13. In any generation, of the q A' -bearers, s will die without reproducing and (1-s) will
- 14. We can now define the total cost of natural selection as: C = S [sq /(1
- 15. J.B.S. Haldane 1892-1964 Haldane was born in Oxford to John Scott Haldane, a physiologist, scientist, a
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